The ratio of the area of the rectangle bounding the seed to the area of the rectangle bounded by the seed
At least three inflorescences for each plant were bagged before anthesis and manually crossed to another individual from the same population of origin. Seeds were let to dry on the plant, collected, cleaned and store in an hermetically sealed box containing silica gel. Eigth seeds from each plant were imaged using a high resolution scanner, at a resolution of 2400 dpi. A size and colour standard was included in each picture. Images were analyed using Tomato Analyzer (Rodrigues. et al. 2010); object boundaries in Tomato Analyzer were manually adjusted to exclude seed hair.