The ratio of the area of the rectangle bounding the leaf to the area of the rectangle bounded by the leaf
The youngest, fully developed leaf from the primary stem was collected at anthesis and scanned on a flatbed scanner, at a resolution of 300 dpi. A size and colour standard was included in each picture. For morphometric measurements, colours were inverted in Adobe Photoshop to increase contrast. Images were analyed using Tomato Analyzer (Rodrigues. et al. 2010); leaf boundaries were manually adjusted when necessary.